A Revision of the Genus Acromastigum by Alexander W. Evans

By Alexander W. Evans

The writer's genusAcromastigum( I zero) illustrates terminal branching from either lateral and ventral segments and is the single genus of the Hepaticae with which the latter kind of branching has been linked. The branches coming up during this unprecedented demeanour are flagelliform in personality, and every indicates on the base a slender and incomplete underleaf. This represents the a part of the section that was once no longer inquisitive about the formation of the department. within the writer's dialogue of branching within the leafy Hepaticae (II, p. 23), terminal branching from ventral segments is well unique because the "Acromastigztm" variety, and its precise good points are mentioned. within the genus Bazzania, or M astigobryum because it is usually referred to as, the ventral flagelliform branches are intercalary in beginning and come up within the axils of underleaves. with the exception of this significant distinction the genera Acromastigum and Bazzania have a lot in universal, and the one species of Acromastigum which has to this point been well-known used to be origi nally referred, with a few doubt, to the genus M astigobryum. This species, that's referred to now as A. integrifolium (Aust.) Evans, is outwardly limited to Hawaii.

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The basal portion for about twothirds the length of the leaf is evenly convex, and the ventral margin appears in profile view, when seen from below. In the outer third of the lea:f the convexity is still marked in the dorsal part. A narrow strip along the ventral side, however, becomes plane or nearly so, and a distinct fold can be demonstrated between the convex portion and the plane portion. When an attached leaf is examined from below the plane portion shows particularly well, and it is evident that the transition between the convex basal portion of the ventral side and the plane portion is gradual.

Acute sinus. They are very Evans. A. 15 x 50. B. Part ofplant,ventra lview, mm. 15 nun. X 50. C. Cross-section of branch, x 225. D. Cross-section of branch, showing an underleaf and the two divisions of a side-leaf, x 100. 1::, F. Leaves, X 225. G, H. Und e rleaves, X 225. I. Leaf of flagelliform branch, X 225. The figures were drawn from the type-material. 08 --0. 12 mm. in width at the apex. Both divisions are triangular and acute, and the apex is tipped either with a single cell or with a row of two cells.

S, C) not infrequently show the characteristic number of cortical cells, although the leaves are imbricated. Those of the dorsi-lateral rows measure 2S-34fL in tangential width by 2S-30 fL in radial width, whereas the corresponding figures for the ventral cortical cells are IS-20 fL by IS fL· The medullary cells average about IS fL in diameter and are therefore scarcely smaller than the ventral cortical cells. The outer walls of the dorsilateral cells are, in most cases, 8-10 fL thick, but those of the ventral cells are rarely more than 4 fL thick.

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